2.) Andren, H. 1994. Effects of Habitat Fragmentation
on Birds and Mammals in Landscapes with Different Proportions of Suitable
Habitat - A Review. Oikos 71(3): 355-366.
The author reviewed the literature to make the generalization that
in bird and mammal populations, connectivity is severely effected by landscape
fragmentation.
3.) Fahrig, L. and G. Merriam. 1994. Conservation
of Fragmented Populations. Conservation Biology 8(1): 50-59.
The authors present evidence that landscape spatial structure is essential
to understanding fragmentation among populations.
4.) Gardner, R.H. and R.V. O'Neill. 1991. Pattern, process, and
predictability: the use of neutral models for landscape analysis. Pages
289-307 in Turner and Gardner (1991).
The authors investigate the effects of percolation thresholds on neutral
landscape models that demonstrate various levels contagion.
5.) Green, D.G. 1994. Connectivity and complexity
in landscapes and ecosystems.
Pacific Conservation Biology , in press. Available from the Internet.
URL: http://www.csu.edu.au/people/dgg/esa93.html
Through investigating the effects of disturbance regimes on species
diversity in simulated landscapes, the author, discovered a phase transition
at critical levels of disturbance. Hence, as disturbance increased
linearly, species diversity would suddenly decrease exponentially.
6.) Gustafson E. J. and R. H. Gardner. 1996. The
effect of landscape heterogeneity on the probability of patch colonization.
Ecology 77(1): 94-107.
The authors use an individual-based dispersal model to measure connectivity
among suitable habitat patches on simulated neutral landscapes.
7.) Gustafson, E. J. and G. R. Parker. 1992. Relationships
between landcover proportion and indices of landscape spatial pattern.
Landscape Ecology 7(2): 101-110.
The authors simulate binary neutral landscapes (forest, non-forest)
to quantify the performance of percolation models with empirical data.
8.) Keitt, T.H., D.L. Urban, and B.T. Milne. 1997. Detecting
critical scales in fragmented landscapes. Conservation Ecology [online]1(1):
4. Available from the Internet. URL: http://www.consecol.org/vol1/iss1/art4
The authors use simulated landscapes to quantify habitat connectivity
at hierarchical scales, and make comparisons with empirical data.
The investigation demonstrated that connectivity among a landscape is scale
dependent.
9.) Milne, B. T. 1992. Spatial Aggregation and Neutral
Models in Fractal Landscapes. American Naturalist 139(1): 32-57.
The author developed a neutral model of species co-occurrence to investigate
the effects of scale on organism parameters (i.e., home range size, dispersal
ability, or speed with which organisms use resources) in fragmented landscapes.
10.) Milne B. T., Johnson A. R., Keitt T. H., Hatfield C. A.,
David J. and P. T. Hraber. 1996. Detection of critical densities
associated with pinon-juniper woodland ecotones. Ecology 77(3): 805-821.
The authors simulated the gradated neutral landscapes to assess ecotonal
effects of encroaching grasslands on pinon-juniper woodlands.
11.) O'Neill, R. V., R. H. Gardner and M. G. Turner. 1992.
A hierarchical neutral model for landscape analysis. Landscape Ecology
7(1): 55-61.
The authors present a methodology for generating hierarchical neutral
landscapes based upon landscape metrics (i.e., contagion).
12.) Palmer, M. W. 1992. The Coexistence of Species
in Fractal Landscapes. American Naturalist. 139(2): 375-397.
The author simulated species interacting across neutral landscapes
with varying fractal dimensions. The investigation demonstrated that
as the fractal dimension (increase in mass) approached its euclidian dimension,
the beta diversity decreased while connectivity increased.
13.) Plotnick, R. E., Gardner, R. H. and R. V. Oneill.
1993. Lacunarity indexes as measures of landscape texture.
Landscape Ecology 8(3): 201-211.
The authors use lacunarity as an index to measure degrees of
texture in 1-3 dimensions of simulated neutral landscapes with graduated
levels of percolation, contagion, and self-similarity.
14.) Turchin Peter. 1996. Fractal analyses of animal movement:
A critique. Ecology 77(7): 2086-2090.
The author utilizes neutral random walk models to determine the
validity of fractal analysis techniques in helping to better understand
animal movement paths. He concludes that the use of the fractal dimension
as a metric to assess movement patterns is weak due to their relatively
poor self-similarity and bias to the small scales on which most animals
move.
15.) With, K. A. 1997. The application of neutral landscape
models in conservation biology. Conservation Biology 11(5): 1069-1080.
The author suggest how concepts from neutral landscape models can be
applied to issues in conservation biology; i.e. metapopulation models.
She stresses that development of a generalized, spatially explicit framework
is vital to the development of individual (species) based modeling.
16.) With, K. A. and T. O. Crist. 1995. Critical
Thresholds in Species Responses to Landscape Structure. Ecology 76(8):
2446-2459.
The authors use landscape based neutral models to investigate changes
in spatial patterns known as critical thresholds. These thresholds
relate to dispersal potential (connectivity) among patches, and are of
importance to better understand the encroachment of habitat loss on threatened
and endangered species.
17.) With, K. A., Gardner, R. H. and Turner, M. G. 1997.
Landscape connectivity and population distributions in heterogeneous environments.
Oikos 78(1): 151-169.
The authors use neutral landscape models to demonstrate that the quality
of landscape connectivity is a function of its percolation threshold and
fractal like self-similarity.
18.) With, K. A and A. W. King. 1997. The use and
misuse of neutral landscape models in ecology. Oikos 79(2): 219-229.
The authors clarify the purpose of neutral landscape models and review
the literature on applications to ecology. They emphasize the application
of neutral models to future studies, rather than theoretical developments.
19.) With, K. A. and A. W. King. 1999. Dispersal
success on fractal landscapes: a consequence of lacunarity thresholds.
Landscape Ecology 14(1): 73-82
The authors use neutral models coupled with landscape metrics (i.e.,
contagion, and lacunarity) to investigate the effects of habitat fragmentation
on dispersal success.
20.) With, K. A. and A. W. King. 1999. Extinction
thresholds for species in fractal landscapes. Conservation Biology
13(2): 314-326.
The authors use neutral models to investigate the effects of habitat
loss and fragmentation on extinction thresholds. Through simulated
fractal landscapes extinction thresholds are developed for species that
are endangered.
Books
Grimmett, Geoffrey. 1989. Percolation. New York: Springer-Verlag,
296p.
Nitecki, M. H. and Antoni Hoffman. 1987. Neutral models in biology. New York: Oxford University Press, 166p.
Stauffer, Dietrich. 1987. Introduction to percolation theory. Philadelphia: Taylor & Francis, 124p.
Turner, M. G. and R. H. Gardner. 1991. Quantitative Methods in Landscape Ecology. New York: Springer-Verlag,
Articles
Gardner, R. H., B. T. Milne, M. G. Turner, and R. V. O'Neill.
1987. Neutral models for the analysis of broad-scale landscape pattern.
Landscape Ecology 1(1): 19-28.
Keitt, T. H. and A. R. Johnson. 1995. Spatial Heterogeneity and Anomalous Kinetics - Emergent Patterns in Diffusion Limited Predatory Prey Interaction. Journal of Theoretical Biology 172(2): 127-139.
Keitt, T. H. 1996. Spectral Representation of Neutral Landscapes.
Working Paper, Santa Fe Institute. Available from the Internet. URL: http://www.santafe.edu/sfi/publications/Working-Papers/96-12-087.ps