Outline, ZOO 3713C, Fall Semester, 2003

 

Introduction

 

The comparative approach as insight into evolution

 

Terms and concepts

·      Ontogeny

·      Embryology

·      Phylogeny

·      Analogy

·      Homology

·      Divergent Evolution

·      Homoplasy

·      Reversal

·      Convergence

·      Parallel Evolution

 

More about homology and analogy

·      How do we establish homology

 

The concept of recapitulation

·      Haeckel’s original proposition

·      von Baer’s Law

·      Biogenetic Law

·      Developmental constraint and integration

·      Phylogenetic constraint

 

Important evolutionary principles

·      Continuity of structure

·      Use and disuse

·      Heterochrony

·      Neoteny

·      Paedomorphosis

·      Hypermorphosis

·      Coexistence and Ancestry

·      Adaptation through natural selection

·      Constancy of environmental change

 

Most evolution is adaptive

Natural Selection

Anagenesis = phyletic transformation

Cladogenesis = splitting = speciation

 

 

The Phylum Chordata

 

Basic chordate characters

·      notochord

·      dorsal hollow nerve chord

·      pharyngeal “gill” slits or clefts

 

“Protochordates”

·      Urochordata - sea squirts, larvaceans, thaliaceans

·      Cephalochordata - amphioxus or lancelet

 

Vertebrata

·      Vertebrate characteristics

·      Vertebrate diversity and classification (details in handout)

·      Review of vertebrate phylogeny and evolution

 

 

Embryology

 

Egg size, cleavage patterns, yolk distribution

 

Origin of embryonic germ layers

·      Ectoderm

·      Endoderm

·      Mesoderm

·      Chordomesoderm

·      Notochord

·      Different mechanisms of “mesoderm” formation

·      Organization of the vertebrate body plan relative to the germ layers

·      Derivatives of the germ layers

 

Skeletal System

 

Exoskeleton

 

Endoskeleton

·      Tissues - notochord, bone, cartilage, dentine, enameloid

 

Cartilage

·      Composition

·      Types

·      Growth

 

Bone

·      Composition

·      Types

·      Growth

·      Joints

 

Organization of the endoskeleton

·      Axial

·      Vertebral Column

·      Ribs and Sternum

·      Skull

·      Visceral Skeleton

·      Appendicular

·      Paired and unpaired appendages

·      Girdles

 

Axial Skeleton

·      Vertebral column

·      Evolution

·      Embryological origin of vertebrae

·      Fossil history of vertebrae

·      Types and diversity of modern vertebrae

·      Regionalization

·      Ribs

·      Primary vs Secondary Ribs

·      Dorsal ribs and ventral (pleural) ribs in fishes

·      Articulation of ribs with vertebrae

 

·      Sternum

·      Origin and variation

·      Articulation with ribs

 

Appendicular Skeleton

·      Origin of paired appendages

·      Paired Fins

·      Lobe fins vs ray fins

·      Organization of the tetrapod limb

·      Modifications of the basic plan

·      Pectoral Girdle

·      Primary Girdle

·      Dermal Girdle

·      Pelvic Girdle

·      Unpaired Appendages

·      Strong convergence - dorsal fins

·      Caudal fin types in fishes

 

Visceral Skeleton

·      No jaws vs jaws

·      Gills vs no gills

·      Generalized gill arch and possible evolution of jaws

·      The splanchnocranium vs the hyobranchium

·      Types of jaw suspension

·      Derivatives of the visceral skeleton

·      Jaws - splanchnocranial components plus dermal skeleton additions

 

Skull

·      Major building blocks

·      Neurocranium

·      Sensory capsules

·      Dermocranium (Dermatocranium)

·      Splanchnocranium

·      Overview of mammalian skull organization

·      Variations on the basic plan

·      Assimilation, fusion

·      Decrease in dermal elements in skull

·      Hypertrophy of dermal jaws

·      Evolution of sound conducting routes

·      Changes in the mandibular suspensirium

·      Dissociation of skull and pectoral girdle

·      Formation of temporal fossae

·      Formation of secondary palate

·      Cranial kinesis

 

 

Muscular System

 

Types of muscle tissue

·      Smooth (Involuntary)

·      Striated (Skeletal, Voluntary)

·      Cardiac

 

Primitive, basic organization of skeletal muscles

 

Embryonic origin of muscle types

·      Epimere - myotomic muscles

·      Hypomere - branchiomeric muscles

 

Axial Muscle Organization

·      Epaxial

·      Hypaxial

 

Appendicular Muscles

·      Primary - Intrinsic

·      Secondary - Extrinsic

 

Integumentary Muscles

·      Types, origins

 

Facial Muscles

·      Types, origins, importance

 

Electric Organs

·      Types, origins, importance

 

 

Integumentary System

 

Basic organization of vertebrate integument

·      Epidermis

·      Dermis

·      Variations on the plan

 

Dermis and its derivatives

·      Primary dermal armor

·      Dermal scales, scale evolution

·      Secondary dermal armor

 

Epidermis and its derivatives

·      Epidermal seal

·      Epidermal growth, shedding

·      Epidermal scales

·      Feathers

·      Hair

·      Nails

·      Horns

·      Glands

 

Skin Pigments

 

 

The Coelom

 

Organization

 

Compartments

 

Stages of evolution

 

 

Digestive System

 

Buccal Cavity

·      Teeth

·      Origin

·      Basic anatomy

·      Number, form, replacement, attachment, modifications

·      Tongue

·      Oral glands

 

Esophagus

·      Crop

 

Stomach

·      Basic organization

·      Modifications

·      Gizzard stomach

·      Ruminate stomach

 

Intestine

·      Regionalization

·      Variations in length

·      Valves

 

Liver

 

Pancreas

 

Respiratory System

 

Olfactory system

·      Choanae

 

Organs of aquatic respiration

·      Internal pharyngeal gills

·      External pharyngeal gills

·      Other

 

Organs of aerial respiration

·      Lungs

·      Other

 

Review contributions of pharynx

 

Evolution of pharyngeal gills

·      Derivatives of the pharynx

 

Structure of internal gills in a shark and bony fish

 

Structure and distribution of external gills

 

Air Bladders vs lungs vs swim bladders

·      Characteristics of lungs

·      Characteristics of swim bladders

·      Intermediate types

 

Fish with pharyngeal gills and lung-like swim bladders

 

Tetrapod lungs

·      Basic structure

·      Basic lung function

·      Unique structure of bird lungs

 

 

Urogenital System

 

How the urinary and reproductive systems are related

 

The Holonephros

 

Glomerular structure

·      Stages of evolution

·      Relationship to coelom

·      Structure and function

 

Structure and function of the metanephros

 

The peritubular capillary network

 

Kidney types in vertebrates

·      Embryonic, transitional, adult

·      Holonephros (Archinephros)

·      Pronephros

·      Opisthonephros

·      Mesonephros

·      Metanephros

 

Osmoregulation in fish

 

Kidney and genital ducts

·      Mullerian ducts, oviducts

·      Holonephric, opisthonephric ducts

·      Wolffian ducts

·      Ureter

·      Ductus deferens

 

Evolution of urogenital ducts

 

Male-female homologies

 

Regionalization of the oviducts

·      Types of uteri in mammals

 

The cloaca

·      Relationship to digestive, urinary, and reproductive systems

 

Urinary Bladder

·      Tubal

·      Cloacal

·      Allantoic

 

Gonads

·      Basic anatomy and function

·      Testicular Displacement

 

Copulatory organs for internal fertilization

·      Gonopodium

·      Hemipenes

·      Claspers

·      Penis

 

Male-female homologies in external genitalia

 

The Cleidoic Egg

 

Problems in vertebrate reproduction

·      Challenges of the terrestrial environment

 

Adaptations of the cleidoic egg

 

Extraembryonic membranes

 

 

Circulatory System

 

Components

·      Vascular system

·      Lymphatic system

·      Hemopoietic organs

·      Blood

 

Structure of arteries, veins, capillaries

 

Hypothetical prevertebrate vascular system

·      Arterial plan

·      Venous plan

 

Heart

·      Location, basic structure

·      Changes during evolution

·      Fish stage

·      Early tetrapod stage

·      Primitive cold-blooded stage

·      Advanced cold-blooded stage

·      Warm-blooded stage

·      Diversity in reptiles

 

Aortic arches

·      Primitive structure, organization

·      Modifications in fishes

·      Modifications in tetrapods

·      Embryological evidence

·      Circulation to external gills

 

Modifications of the primitive venous plan

·      What is a portal system

·      Evolution of the hepatic portal system

·      Evolution of the postcava

·      Formation of the renal portal system

·      Formation of the protopostcava

·      Formation of supracardinals

·      Loss of renal portal system

·      Evolution of the precava

 

 

Nervous System

 

Components

·      Central

·      Peripheral

 

Structure and function of a neuron

 

Nerve fiber types

·      Motor (efferent)

·      Sensory (afferent)

 

Cranial Nerves

·      Numbers, names of cranial nerves

·      Evolution of cranial nerves

 

Spinal Nerves

·      Structure of spinal nerves

·      Motor and sensory pathways

·      Associated ganglia

·      Nerve plexi

·      Evolution of pathways

 

Anatomy of the spinal cord

 

Simple reflex arc

 

Autonomic nervous system

·      Symapthetic

·      Function

·      Nerves involved

·      Preganglionic fibers

·      Postganglionic fibers

·      Collateral ganglia

·      Parasympathetic

·      Function

·      Nerves involved

·      Preganglionic fibers

·      Postganglionic fibers

·      Ganglia

 

Meninges of the CNS

 

Brain

·      Basic organization

·      Ventricles

·      Ontogenetic/phylogenetic development

·      Major landmarks in the brain

·      Structure

·      Function